From mboxrd@z Thu Jan  1 00:00:00 1970
From: Mark A. Hershberger <mah@everybody.org>
Subject: Re: Bug: check that org-element--cache-sync-keys is non-nil before
 clrhash [8.3.4 (8.3.4-42-gae73c7-elpa @
 /home/mah/.emacs.d/elpa/org-20160425/)]
Date: Sun, 22 May 2016 14:16:45 -0400
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To: Nicolas Goaziou <mail@nicolasgoaziou.fr>
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> Could you investigate about how this happens in your case?

I removed [gnus-alias-use-identity](https://www.emacswiki.org/emacs/GnusAlias) and the problem went away.  I'm guessing it is because some sort of race condition when I reply to a message.  I would gues that org-mode is invoked when the signature files are inserted into the message.